- Received May 13, 2013.
- Accepted June 11, 2013.
- Published online July 17, 2013.
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© 2013 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0/, which permits unrestricted use, provided the original author and source are credited.
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- Scott D. Sampson1,2,3⇑,
- Eric K. Lund2,3,4,
- Mark A. Loewen2,3,
- Andrew A. Farke5 and
- Katherine E. Clayton3
- 1Denver Museum of Nature and Science, 2001 Colorado Boulevard, Denver, CO 80205, USA
- 2Department of Geology and Geophysics, University of Utah, 115 S 1460 East, Salt Lake City, UT 84112, USA
- 3Natural History Museum of Utah, University of Utah, 301 Wakara Way, Salt Lake City, UT 84108, USA
- 4Department of Biomedical Sciences, Ohio University, Heritage College of Osteopathic Medicine, Athens, OH 45701, USA
- 5Raymond M. Alf Museum of Paleontology, 1175 West Baseline Road, Claremont, CA 91711, USA