- Received January 6, 2014.
- Accepted February 12, 2014.
- Published online March 11, 2014.
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© 2014 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0/, which permits unrestricted use, provided the original author and source are credited.
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- 1Wildlife Biology Program, Department of Ecosystem and Conservation Sciences, University of Montana, Missoula, MT 59812, USA
- 2US Geological Survey, Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT 59812, USA