- Received January 17, 2014.
- Accepted April 2, 2014.
- Published online April 30, 2014.
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© 2014 The Author(s) Published by the Royal Society. All rights reserved.
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- 1National Oceanic and Atmospheric Administration (NOAA), Pacific Marine Environmental Laboratory (PMEL), 7600 Sand Point Way NE, Seattle, WA 98115, USA
- 2Conservation Biology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration (NOAA), 2725 Montlake Boulevard East, Seattle, WA 98112, USA
- 3NOAA NMFS NW Fisheries Science Center, 2030 SE Marine Science Drive, Newport, OR 97365, USA
- 4Oregon State University, Cooperative Institute for Marine Resources Studies, Hatfield Marine Science Center, 2030 SE Marine Science Drive, Newport, OR 97365, USA
- 5College of Earth, Ocean, and Atmospheric Sciences, Oregon State University, Corvallis, OR 97331, USA