- Received July 7, 2014.
- Accepted October 7, 2014.
- Published online November 5, 2014.
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© 2014 The Author(s) Published by the Royal Society. All rights reserved.
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- 1Evolutionary Ecology Unit, Department of Biology, Lund University, Lund 223 62, Sweden
- 2Centre for Ecological and Evolutionary Synthesis (CEES), Department of Biology, Oslo University, Oslo, Norway