- Received June 20, 2014.
- Accepted October 16, 2014.
- Published online November 12, 2014.
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© 2014 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.
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- 1Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT, USA
- 2Department of Integrative Biology, Oregon State University, Corvallis, OR, USA
- 3Department of Biology, Butler University, Indianapolis, IN, USA
- 4Department of Ecology and Evolutionary Biology, Cornell University, Ithaca, NY, USA
- 5Department of Biological Sciences, National University of Singapore, Singapore
- 6NUS-Yale College, Singapore