- Received July 9, 2015.
- Accepted September 4, 2015.
- Published online September 30, 2015.
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© 2015 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.
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- 1Department of Animal and Plant Sciences, University of Sheffield, Western Bank, Sheffield S10 2TN, UK
- 2Animal Ecology, Department of Ecology and Genetics, Uppsala University, Uppsala 752 36, Sweden
- 3Centro Nacional del Medio Ambiente, Universidad de Chile, Avenida Larrain 9975, La Reina, Santiago, Chile