- Received August 10, 2009.
- Accepted October 12, 2009.
- Published online November 11, 2009.
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© 2009 The Royal Society
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- 1Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Johannesburg 2050, South Africa
- 2Department of Biological Sciences, Western Illinois University, Macomb, IL 61455, USA
- 3Council for Geoscience, Pretoria 0001, South Africa
- 4Zoology Department, University of Cape Town, Private Bag X3, Rhodes Gift 7700, South Africa
- *Author for correspondence ( ).