- Received February 16, 2011.
- Accepted March 25, 2011.
- Published online April 13, 2011.
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- 1Department of Entomology, Cornell University, Ithaca, NY 14853, USA
- 2Agricultural Research Council, Private Bag X134, Queenswood 0121, South Africa
- 3School of Biological and Conservation Sciences, University of KwaZulu-Natal, Private Bag X01, Scottsville, Pietermaritzburg 3209, South Africa
- 4Laboratory of Evolutionary Entomology, Institute of Biology, University of Neuchatel, Emile-Argand 11, 2000 Neuchatel, Switzerland
- *↵ Author for correspondence ( ).